The Evolution of Flight
CHAPTER 13: UPDATES
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OCTOBER 2005. NEW THOUGHTS ON PTEROSAUR FLIGHT (p. 181)
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FEBRUARY 2005. Gliding Ants (p. 179)
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New flight model for pterosaurs. Basically, the novelty is in proposing a flap effect for a strip of tissue along the leading edge of the wing, manipulated by the pteroid bone. The idea was tested in a wind tunnel. The "flap" would have been used mainly on take-off and landing, but could have been used to do barrel-rolls in flight (if needed!). This function would be analogous to the flaps on an airplane (aeroplane for Brits). And all pterosaurs had it. The paper was in the Proceedings of the Royal Society of London, October 2005. I think it is a classic. The experiments were done in Ellington's lab in Cambridge, so you can be sure about them. What they have done is to take away any lingering doubt there may have been that pterosaurs could take off and land effectively.
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Gliding ants. Here's something else to add to the list of animals that have evolved gliding. Press release from UC Berkeley, February 2005. Astounding: but there are probably lots more of them if anyone looked.
CHAPTER 13: NOTES AND WEB LINKS
Lecture Notes on weed seed dispersal
Flight in Insects
P. 178. Swimming (and skimming) as a preadaptation
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Discover feature on Jarmila Kukalova-Peck's work, among other things: from 2000.
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Movies showing the relationship between gills and wings in some living mayflies and stoneflies. From James Marden's lab at Penn State.
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James Marden's research on stoneflies
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Class exercise on evolution of insect flight by skimming stoneflies
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Summary of Marden and Kramer, 1995
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A new cladistic analysis of living stoneflies, based on molecular evidence as well as wing structure and flight style, shows that the most "basal" of living stoneflies are surface skimmers. This suggests, though it can't prove, the stonefly flight began with skimming. The implication, of course, is that the same might apply to all insect flight. Reference: Thomas, M. A., et al. 2000. Molecular phylogenetic analysis of evolutionary trends in stonefly wing structure and locomotor behavior. Proceedings of the National Academy of Sciences, published online before print November 14, 2000.
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Links to videos of skimming stoneflies, from James Marden's lab at Penn State.
Flight in Vertebrates
UC Berkeley's pages on vertebrate flight: the subsections include pages on most of the sections below.
P. 179. Parachuting Vertebrates
P. 179. Early Gliding Vertebrates
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Coelurosauravus.
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Icarosaurus
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Sharovipteryx, a Late Triassic gliding lepidosaur
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Longisquama, a Late Triassic gliding lepidosaur
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Longisquama, including a photograph of the real specimen.
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Longisquama had scales, not feathers. From BBC News OnLine. This is a response to an earlier paper in June 2000. Some Web pages from that date:
Pterosaurs
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Introduction to Pterosaurs from UC Berkeley
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Pterosaurian flight from the UC Berkeley site: you can bet that Kevin Padian had a lot of input into this one!
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A pterosaur site by an enthusiast
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A foot fossil of the small early pterosaur Dimorphodon suggests that it could not have run: Press release, 1998.
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This site includes a nice reconstruction of the Jurassic pterosaur Sordes pilosus. It is alleged to have been hairy, though that is disputed. If so, then pterosaurs (or some pterosaurs) were warm-blooded.
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A new pterosaur (from the Cretaceous of Brazil) that fished by skimming the water.
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The largest pterosaur was Quetzalcoatlus may have been the ecological equivalent of a gigantic heron, fishing in inland lakes for frogs or turtles or arthropods such as crayfish from shallow water.
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The brilliant paper by Larry Witmer and colleagues on pterosaur brains, balance, and behavior, published October 2003:
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Pterosaur nests have been found occasionally, but a major nesting area has been discovered in Chile.
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A pterosaur embryo inside a pterosaur egg. Originally from Nature news service, June 10, 2004.
Birds
Birds are much underrated in terms of intelligence: "bird-brain" is a term of abuse in American vernacular. The reality is that (modern) birds are very clever, and it may be that the cunning of dinosaurs was one of things that Michael Crichton got right in Jurassic Park.
Emery, N. J., and N. S. Clayton 2004. The mentality of crows: convergent evolution of intelligence in corvids and apes. Science 306: 1903-1907. When you read it, you will find that corvids (crows) are on the same level as primates in terms of intelligence and learning (tool use, for example).
How clever are crows?
The Solnhofen Limestone
Home of Archaeopteryx, the earliest bird, and many beautiful pterosaur fossils.
Early Birds, including Archaeopteryx
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Fossil record of birds
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The earliest known bird Archaeopteryx
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"All about Archaeopteryx"
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The Archaeopteryx Web Site: stories, pictures, animations, links.....
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Lithograph of Archaeopteryx published by Richard Owen in 1863
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Archaeopteryx on the EvoWiki site.
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CT scan of an Archaeopteryx brain. The paper and a commentary by Witmer are in Nature, so won't be freely available on the Web. Dominguez Alonso, P., et al. 2004. The avian nature of the brain and inner ear of Archaeopteryx. Nature 430: 666669, and commentary by Larry Witmer, pp. 619620. This is a very fine study. The brain is very bird-like, though as you might imagine is primitive bird-like. No matter what the authors hint, this is not proof that Archaeopteryx flew (and Witmer says that too). The study does show clearly that Archaeopteryx is at least very close to the origin of flight. I have written a longer comment here.
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Press release from Tim Rowe's lab at the University of Texas
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Originally from Nature news service
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News Scientist.
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BBC News OnLine
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National Geographic news
The Origin of Feathers
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Evolution of feathers essay on EvoWiki. Its weakness is that it ignores the dramatic array of evidence for feathers in non-flying theropods (as laid out in this EvoWiki article, for example), so the discussion is bird-oriented and therefore the conclusions are suspect. For example, they don't like the display hypothesis :)
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How feathers form in chickens.Press release, October 31, 2002. The hope is (and it is only a hope) that the sequence seen in chickens is also the evolutionary sequence that took place 150 million years ago. Not guaranteed!
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Many theropod dinosaurs had feathers.. That means that feathers did not evolve for flight. So how/why did they evolve? I naturally like an idea that I developed jointly with my colleague Jere Lipps. Here, for historical interest, is what Jere Lipps and I wrote in 1982.
The Origin of Powered Flight in Birds
The Arboreal Hypothesis
I have to say that I think the arboreal hypothesis is so weak that it's not worth writing about. See a brief note that I wrote.
The Display and Fighting Hypothesis
The WAIR hypothesis
was published in January 2003. Here is the paper, by Ken Dial. (Thank you, Science!). Stories at the time:
Here's my take. The observations on living birds are fascinating, but the application to the origin of bird flight is wrong, in my opinion. I wrote a very brief note to Science about it, but it was not published: Letter to Science, submitted February 2003, rejected March 2003.
Did Archaeopteryx Fly?
It is fair to say that the majority opinion is that Archaeopteryx could fly. . (Article on EvoWiki.)
I suspect, however, that Archaeopteryx could not fly. Part of the problem as I see it, as I say in the book, is that Archaeopteryx could not have had an effective upstroke without the supracoracoideus system.
In May 1999, Nature published a paper by Burgers and Chiappe, which claimed that Archaeopteryx ran well, and was able to take off by flapping its wings and running. I have problems with this paper, and will discuss them here eventually. In a nutshell, I suspect that Burgers used equations from living birds that fly to do his calculations on Archaeopteryx. In other words, he assumed his answer first. But he avoids saying that, even in the small print of the footnotes... Stay tuned. Here is the paper, rightly or wrongly posted on the Web, but not by Nature. And here is a story from the San Francisco Chronicle.
For my latest take on the lack of flight in Archaeopteryx, see this mini-essay from August 2004.
Earlier Bird Evolution?
New Cretaceous Birds
New discoveries of Cretaceous birds and close relatives:
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Jeholornis, a new primitive fossil bird from China, with seeds preserved in its gut.
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Protarchaeopteryx from China.
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Rahonavis is a small flying Cretaceous bird from Madagascar, but it has a slashing claw much like its distant coelurosaur cousins. The Science paper.
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Feature on the fossil deposits of Liaoning, China, from BBC News OnLine.
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Confuciusornis, an Early Cretaceous bird from China. (It has DISPLAY feathers!). See also the next two images, here and here
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Protopteryx, a new very early bird from China.
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A feathered bird chick from the Lower Cretaceous of China. It is tucked up as if it died inside the egg at a very late stage, and it's strong-boned and feathered. If it's feathered at this stage, it would have hatched as a precocial chick. That means (to me) that the bird was NOT nesting in a tree, but on the ground. Yet the artist, Zongda Zhang, has portrayed the birds as hatching in a nest in a tree! I can't think of a precocial chick today that hatches from a nest in a tree. Maybe there's some inscrutable logic that requires (maybe even permits) such an interpretation, but I don't see it. The Beijing group working on early birds seem to be fixated on an arboreal origin for flight (so maybe for birds?), and this may be an offshoot of that viewpoint, in the face of the evidence. Or maybe I am biased: I am on record in the book as saying the arboreal origin of flight is nutty (to paraphrase). Anyway, read the blurb, read the paper (which is in Science, October 2004), and decide for yourself.
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Eoalulavis from Spain has evidence of excellent slow-flight control. Story originally from Natural History.
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Asparavis, a new Cretaceous bird from Mongolia: it is one of the earliest birds that lies in the direct ancestry of surviving birds today.
Press release from Yale University; the same story is
here.
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Mononykus, from the Late Cretaceous of Mongolia, is an example of an early flightless terrestrial bird. Mononykus and its close relative Shuvuuia
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A Web essay on Ichthyornis from the Oceans of Kansas Web site. Updated 2005.
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A Web essay on Hesperornis from the Oceans of kansas Web site. Updated 2005.
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A reconstruction of the flightless fishing bird Baptornis from National Geographic.
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Cretaceous birds already had bird lice -- now there's some rapid evolution for you! (Link broken as Nature re-structured its news service, July 2004).
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A Cretaceous duck? The paper is in Nature, so won't be on the Web. This is part of a huge argument about how many lineages of "modern" birds were present before the K-T boundary. The fossil is from the latest Cretaceous, but it is from Antarctica, so it won't be easy to try to collect more. The specimen is either an "unidentifiable bundle of bones", or has bones with unquestioned affinity to the Anatidae, the family to which modern ducks belong. I'm not willing even to guess who is right. It would be a surprise to me if the specimen really is an anatid, but stranger things have turned up.
Cenozoic Birds
The Largest Flying Birds
Bats
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Flight in bats
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Tree of Life Website on Bats, with links at the end of the section.
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A bat that eats birds: BBC News OnLine, August 2001.
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How vampire bats take off. (They jump high enough to get in a couple of powerful wing beats. This is not likely to have been the way that bats originally evolved flight.)
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How bats invented butterflies. BBC News OnLine, January 2000. Some moths may have evolved into day-flying butterflies to avoid nocturnal bat predation. (No evidence for this story, but it sounds plausible.)
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Some bats have evolved ultraviolet vision: stories from October 2003. Press release about a new paper in Nature. Here is another version of the release with a picture of the bat. Mammals lost color vision: only primates as a major clade re-evolved it. This bat from tropical America searches for nectar in the dark, and is also color-blind. What it has done is to "push" its black-and-white vision into the ultraviolet (the flowers it visits bloom at night, and their petals reflect whatever little light there is, with very high reflections at ultraviolet wavelengths). This looks like impressive co-evolution to me. We will have to wait for more studies on more nectar-sipping bats to see how common it is. It is a very laborious business to do the experiments, however. The paper is at Nature 425: 612-614, but of course Nature does not make its contents universally available on the Web.
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More insight into bat evolution: early and rapid, along with sonar, and probably in the Eocene. The paper and a commentary are in Science, January 2005.
The reference list for Chapter 14, 4th Edition, with associated Web links
Page last updated, October 23, 2005.
Links last checked October 4, 2005.
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